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The 5.56 NATO round was first developed for the battle tested M16 used in Vietnam, and various incantations across the world, to this very day. The 5.56 barrel chamber is mil-spec, allowing more variability in chamber cutting and ammo manufacturing. Why? When you contract manufacturing across the globe, different machines create a lot of variability in machining and ammo. You need to be able to slap a Spanish made barrel on a Colt lower and feed it Russian made ammo from a Chinese magazine, all without a hiccup. Although this is a great scenario for global warfare, it can be troublesome for those wanting to chamber cheap 5.56 high-pressure ammo in their crisp new 223 Rem.
The 223 Rem, while the same caliber as the 5.56, follows SAAMI specifications, meaning it is standardized. If you're going to advertise a 223 Rem, then the barrel chamber and the ammo are different.
Chambering: You can chamber either ammo in either rifle, but because of higher pressures in 5.56 ammo, you could damage your 223 Rem rifle if you use it. Shooting 223 Rem ammo in a 5.56 chamber is fine, but accuracy may degrade.
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223 Wylde to the Rescue: The 223 Wylde is not a new caliber, it is a chambering that accommodates 5.56 ammo pressures with 223 Rem chamber specs, and allows use of either type ammo with the best each has to offer alone. Simply put, a 223 Wylde chamber is used on 223 caliber rifle barrels to allow them to safely fire either 323 Rem or 5.56 NATO ammunition.
Wylde's hybrid chamber was designed to exploit the accuracy advantages of the .223 Remington chambering without problems concerning over pressure or compromising the functional reliability of semi-auto firearms like the AR15 family when using 5.56 NATO military ammunition. Coincidentally, it shoots the relatively long and heavy 80-grain bullets commonly used in the Sport Rifle Competition very well and is one of the preferred chambers for that use.The Wylde chamber is used by rifle manufacturers who sell "National Match" configuration AR15 rifles, barrels, and upper receivers.